(AP) - Joe Paterno, the longtime Penn State coach who won more games than anyone in major college football but was fired amid a child sex abuse scandal that scarred his reputation for winning with integrity, died Sunday. He was 85.

His family released a statement Sunday morning to announce his death.

"He died as he lived," the statement said. "He fought hard until the end, stayed positive, thought only of others and constantly reminded everyone of how blessed his life had been. His ambitions were far reaching, but he never believed he had to leave this Happy Valley to achieve them. He was a man devoted to his family, his university, his players and his community."

Paterno built his program on the credo "Success with Honor," and he found both. The man known as "JoePa" won 409 games and took the Nittany Lions to 37 bowl games and two national championships. More than 250 of the players he coached went on to the NFL.

"He will go down as the greatest football coach in the history of the game," Ohio State coach Urban Meyer said after his former team, the Florida Gators, beat Penn State 37-24 in the 2011 Outback Bowl.

Paterno's son Scott said on Nov. 18 that his father was being treated for lung cancer. The cancer was diagnosed during a follow-up visit for a bronchial illness. A few weeks after that revelation, Paterno also broke his pelvis after a fall but did not need surgery.

Paterno had been in the hospital since Jan. 13 for observation for what his family had called minor complications from his cancer treatments. Not long before that, he conducted his only interview since losing his job, with The Washington Post. Paterno was described as frail then, speaking mostly in a whisper and wearing a wig. The second half of the two-day interview was conducted at his bedside.

"As the last 61 years have shown, Joe made an incredible impact," said the statement from the family. "That impact has been felt and appreciated by our family in the form of thousands of letters and well wishes along with countless acts of kindness from people whose lives he touched. It is evident also in the thousands of successful student athletes who have gone on to multiply that impact as they spread out across the country."

The final days of Paterno's Penn State career were easily the toughest in his 61 years with the university and 46 seasons as head football coach.

It was because Paterno was a such a sainted figure - more memorable than any of his players and one of the best-known coaches in all of sports - that his downfall was so startling. During one breathtaking week in early November, Paterno was engulfed by a scandal and forced from his job, because he failed to go to the police in 2002 when told a young boy was molested inside the football complex.

"I didn't know which way to go...and rather than get in there and make a mistake," he said in the Post interview.

Jerry Sandusky, the former defensive coordinator expected to succeed Paterno before retiring in 1999, was charged with sexually assaulting 10 boys over 15 years. Two university officials stepped down after they were charged with perjury following a grand jury investigation of Sandusky. But attention quickly focused on an alleged rape that took place in a shower in the football building, witnessed by Mike McQueary, a graduate assistant at the time.

McQueary testified that he had seen Sandusky attacking the child and that he had told Paterno, who waited a day before alerting school authorities. Police were never called and the state's top cop later said Paterno failed to execute his moral responsibility by not contacting police.